Brendan Doggett and Scott Boland opened the bowling for the Indigenous XI in 2018 during 150 year commemorations of the 1868 Aboriginal cricket team's tour of England.
That 19th century team was the first ever Australian sporting outfit to head overseas.
More than a century and a half on, two First Nations men will feature in the Test team for the first time following confirmation of Doggett's selection in Australia's XI for the first Ashes Test, starting Friday.
Stand-in captain Steve Smith named Australia's XI for Perth on Thursday.
Northern Territory-raised Tasmania opener Jake Weatherald will also debut with Cameron Green awarded the sole allrounder position, with Beau Webster missing out.
It's the first time Australia have named two debutants in a Test since 2019.
Usman Khawaja will open, with hot domestic form earning Marnus Labuschagne a recall to bat at three.
Smith, Travis Head, Green and 'keeper Alex Carey make up the remaining batters.
England named their 12-player squad on Wednesday, with Jofra Archer and Mark Wood set to lead their pace attack.
A shoulder injury had previously kept England skipper Ben Stokes in doubt, with English all-time leading Test run scorer Joe Root still chasing his maiden Ashes century away from home.
For Australia, Doggett and Boland will partner with Mitchell Starc as the front-line pace trio, with veteran offspinner Nathan Lyon also named.
A back injury to Pat Cummins all but confirmed Boland's inclusion for the start of the series a month ago, with more a recent hamstring strain to Josh Hazlewood opening the door for a historic debutant in Doggett.
The South Australia quick - who has been in career-best form to start the domestic red ball summer - has been a recent touring member with the Test team after first entered the squad seven years ago without breaking into the playing XI.
"He brings a real skill, a fast outswing bowler," Scott Boland said of Doggett earlier this week.
"He's someone who can bowl really long spells.
"He tends to bowl 20 or 25 in a day, (South Australia) push their quicks a little bit more than what some other teams do."
Doggett will become the fifth Indigenous cricketer to receive a baggy green on Friday after men's players Jason Gillespie and Boland, current women's star Ash Gardner and mid-20th century trailblazer Faith Thomas.
The Worimi man's addition is also being framed as one of the feel-good stories in recent years.
A relative unknown until his into his 20s, the Toowoomba-raised quick who was working as a carpenter didn't come through state high performance pathways, instead earning domestic call ups out of local cricket.
Doggett's move to Adelaide ahead of the 2021-22 summer bolstered his CV, helping break South Australia's 29-year Shield drought earlier this year. Previously he won a final with Queensland four years earlier.
Doggett's 11-140, against his former state, are the best-ever figures by a bowler in a Sheffield Shield final.
"I don't know how ready you can be for Test cricket," Doggett said earlier this week, reflecting on his journey.
"But the last 18 months, to two years, has easily been the most successful I've been in terms of numbers (of wickets), but also just confidence in my body, confidence in my game."
He's taken 13 wickets at 14.69 across two matches to start the current season, including two 5-fas.
Boland will feature in his third Ashes series after exploding onto the scene with a fairytale debut four years ago, and early inclusions in 2023's series in England.
The Gulidjan seamer has played 14 Tests, taking 62 wickets at 16.53 after holding his spot from Australia's tour of the West Indies earlier this year.
